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Originally received: July 26, 2016. Final revision: January 3, 2017. Accepted: January 3, 2017. Available online: February 22, 2017. Manuscript no. 2016-1558. 1 State Key Laboratory of Ophthalmology, Zhongshan Ophthalmic Center, Sun Yat-sen University, Guangzhou, China. 2 Department of Radiology, The Affiliated Brain Hospital of Guangzhou Medical University (Guangzhou Huiai Hospital), Guangzhou, China. 3 Research School of Biology, Australian National University, Canberra, ACT, Australia. 4 Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, East Melbourne, Australia. *Xinxing Guo and Ou Xiao made equal contributions to this study. Financial Disclosure(s): The author(s) have no proprietary or commercial interest in any materials discussed in this article. Supported by the Fundamental Research Funds of the State Key Laboratory, National Natural Science Foundation of China (81420108008 and 81271037), Science and Technology Planning Project of Guangdong Province (2013B20400003), and a grant from the Brien Holden Vision Institute. Author Contributions: Conception and design: Guo, Chen, Chen, Morgan, He Data collection: Xiao, Chen, Wu, Chen Analysis and interpretation: Guo, Xiao, Chen, Morgan, He Obtained funding: Not applicable Overall responsibility: Guo, Xiao, Morgan, He