LAUSR

Abstract This study analyzes in detail the beds bearing Rhizocorallium in the Upper Triassic Nayband Formation (Tabas Block, Central Iran) and shows their potential in interpreting depositional settings, ecological features and sequence stratigraphy. An integrated approach combining ichnological, sedimentological and stratigraphic methods has significantly enhanced our understanding of the changes in the spatial distribution style and morphological differences among the same ichnospecies of Rhizocorallium . The Upper Triassic Rhizocorallium assemblages from the Nayband Formation reveal significant spatial variations in the orientation and dimensions (i.e., length, width, limb diameter), the tiering pattern, and the colonization style throughout systems tracts and at erosional discontinuities or omission surfaces. Based on the wide variety of forms and orientation, numerous morphotypes of Rhizocorallium are grouped into two main types, i.e., those that are mostly facies controlled and those that appear abundantly in a confined stratigraphic interval and are facies independent. In this respect, three paleoichnocoenoses or habitats for the rhizocorallid producers could be differentiated according to the burrow morphology and the nature and consistency of the substrate: (1) stable habitat, represented by soft substrates including horizontal, long, straight or slightly sinuous, spreite-bearing, U-shaped protrusive burrows of Rhizocorallium irregulare (=  R. commune var. irregulare ). This habitat resulted from processes operating in settings that are characterized by soft substrates, reduced sedimentation rates, and abundant food supply, typical of low- to moderate-energy, fully marine conditions; (2) unstable, physically-controlled habitat, represented by a suite of shifting substrates with low- to high-energy settings, including short, U-shaped protrusive spreiten-burrows of R. jenense (Type 1) . In this habitat alternating and contrasting energy conditions did exist due to repeated storm events that were the main controlling factor in the distribution and preservation of Rhizocorallium ; (3) substrate-controlled habitat represented by a suite of stiff-to-firm substrates including oblique to vertical protrusive or retrusive spreiten U-shaped burrows of R. jenense (Type 2) . The erosional discontinuities or omission surfaces related to this habitat include parasequence-bounding transgressive surfaces of erosion (TSE) and parasequence-bounding flooding surfaces (FS) and are mainly associated with the Rhizocorallium -dominated Glossifungites ichnofacies. Complementary ichnological information indicates that the shift in the orientation and the morphological features of rhizocorallid burrows point to broad changes of environmental parameters, which are, in turn, controlled by the rate of sea-level changes.